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Volume 18, Supplementto Number 3 15 September1998 JOURNALof VERTEBRATE PALEONTOLOGY ABSTRACTSOF PAPERS FIFTY-EIGHTHANNUAL MEETING SOCIETYOF VERTEBRATEPALEONTOLOGY SNOWBIRDSKI AND SUMMERRESORT SNOWBIRD.UTAH SEPTEMBER 3O-OCTOBER 3, 1998 SOCIETY OF VERTEBRATE PALEONTOLOGY rssN 0272-4634 CIII and CIV. subsequent anatomical differentiation reUects a fiame shift in the developntental processes acting on the digit anlagen in later ontogeny. such that C I I i s t r a n s f o r m e d i n t o a t h u m b . C I I I i n r o a n i n d e x f i n g e r .a n d C I I I i n t o a m i d d l e f i n g e r . P r e l i m i n a r y r e s u l t s f r o m b x p e r i m e n t a l J y - i n d u c e trl n d n a t u r a l l v - o c c u r r i n g variation in archosaur digit homology provide tests tbr this hypornesrs. INFORMATION ON DINOSAUR RESPIRATORY PHYSIOLOGY AND THE STRUCTURE OF THE THEROPOD LUNG GEIST. Nicholas R.. Zoology Department. Oregon Srate Univcrsiry. 3029 C o r d l e y H a l l . C o r v a l l i s ,O R , 9 7 3 3 t - 2 9 t 4 . U S A : H I L L E N I L J S .W i i l e m J . . l ) e p a r m e n l o f B i o l o g y . C o l i e g e O f C h a r l e s r o n .6 6 G e o r g e S t . . C h a r l e s t o n .S C . 2 9 4 2 4 - ( X X ) 1 U . SA. T h e p r e s e n c eo f n a s a l r e s p i r a t o r y t u r b i n a t e s i n m a n t m a l s a n d b i r d s i s c a u s a l l y _ l i n k e d t o e l e v a r e dr o u t i n e r a t e s o l v e n t i l a t i o n a n d h i g h r e s t i n g l e v e l s o l ' o x y g e n c o n s u m p t i o n . A b s e n c e o f r e s p i r a r o r y t u r b i n a r e si n e x t i n c t r e t r a p o d si s a l i k i l y i n d e x o f l o w v e n r i l a t o r y r a r e s a s s ( ) c i r t e dw i r h e c t o t h e r n r i cm e r a b o l i c p h y s i o l o g y . CT-scansindicate thar the relativcly narrow nasal cavities ol- sevcral genera-ol. Late cretaceous theropods were inadecluatet. have housed respiratory (urbinates. New data on the cranial morphology of several theropod. sauropoci.ancl ornithis_ chian dinosaurs confirm the widespread absence of respiratory turhinates in these taxa as well. A d d i t i o n a l l y . ' t h e s t r u c t u r e o f t h e r e c e n t l y c l e s c r i b e dI t a l i a n m a n i r a p t o r a nt h e _ r o p o d " l t l p l o n r : r p r o v i d e s c o m p e l l i n g n e w e v i d e n c e f b r t h e e x i s t e n c eo f a h e p a t i c _ p i s t o n r l i a p h r a r n t i n t h e r o p o d d i n o s l u r s . S p c ' c i l i c a l l v .i n , S c r T r i o r r rr, h _e r,rrorphol_ o g v a n d o r i e n t a l i o n o f t h e I i v e r . t r a c h e . i .a n c j c o l o l a r e c l o s c l v c o n t p a r a b l c1 o l h e i i n a r ( i l r v o l - t h e s c l c a t u r e s i n l i v i n - u c r o c o d i l i a n s . i i n d c o n t r i r s ts h a r P l l r r , i t h t h e i r i l r r a n - s c ' r c n ti n h i r d s . R e l i a n c e < t n a h e p a l i c p i s t o n n t c c h a n i s n t p r i t b a b l v c o n _ s t r a i n e dl h c r o p o d s f r o m d e l e l o p i n g a n a r i a n - s t 1 , l el u n g / a i r s a c r e \ p i r J k ) t , \ s \ s ! e n t . T A P I ' I o N O M Y O F T I I R E E C E N ] ' ] ? ( . ) . S A L I R UB, O ' NI]I]EDS IN THTJ DINOSAI]R PARK FORMA'fION. AI-BERTA. CANAI)A CETTY. Michael A: EBERTH. David A: BRINKMAN. Donalcill. ancl R Y A N . M i c h a e l . 1 .I i o v a l l ' v r r e l l M u s e u m o l p a l t i e o n t o l o g y .B o x 7 5 0 0 . D r u n r h e l l e r .A B T O J 0 Y 0 . C A N A D A . S y s t c n r a t i cs a n t p l i n g o f t h r e e b o n e b c d s c o n s i s r i n g u l n r o s t e x c l u s i v e l l , o f t h e d i s _ articulated remains of Centnt.suuru5Tpe fiL!.\ was conclucted in the lowcr % of the DinosaLrr Park Formation at Dinosirur provincial park iirnr 1995- 1997. An area o f a t l e a s r I 2 n t : o f e a c h b o n e b e d u ' a s s , v s t e m a t i c a l l ey x c a v a t e da n d m r p p e d . M o r e than 907 of identifiable elements belong to Certtro.stturu.srr7rcrltr.r,with bone d e n s i t r ' r a n g i n r l i r r r n 7 0 - 1 8 0 b o n e s / r n r .c ( , t r ( ) ' u r r u s e l e n r e n t sa r e c o n r o l e t e l v d i s a r t i c u l a t e dt n d f e p r e s e n ta r v i d e r a n g e o l a g e c l a s s e s .i r o m i n l h n r r . a t i u l r . A i l of the honebeds occur in overbank nudstone fitcies ancl rre characterized by very s i n r i l a r t a p h o n o n r i cs i g n a t u r e si n d i c u i v e o i n r u l t i - e v e n th i s t o r i e so f 1 ' l r x x i i n gs. c a r , enging and reworking. prior to final burial. T a p h . n o n r i c i n c l i c a t o r ss L l c hi l s r h e w i n n . r v i r g o l s n r a l l e l e r . e n t s . n r i l c la b r a s i . n o f b o n e s a n d p o o r a l i g n m e n t o l e l e n t e n t sd e m c l n s l r a t et l t a t t h e r e w a s a n r o d e s t ( n o n - c h r n n c l j z e r l td c g r e e o f h v c l r a u l i c r v o r k i n c j n l l u e n c i n g t h e h o n c h c d s . Abra_ s i ( ) n o 1 t l x ) l h n r a r k s a n d b r o k L : nc c l g e si n d i c a t e t h a t s r n r e h v d r a u l i c u ' r k i r g . c c u r r e c l s u b s e q u e n r l yr o s c a v e n g i n g a n d / o r t r a m p l i n g . ' f h e b o n e s c x h i b i t f r e s h b r e a k a s e a n d a n a l m o s t c o n r p l e t el a c k , l s u r l i c i a l w e a t h e r i n g \ \ , h i c l t i s c o l t s i s t e n t L r f r e l a t i r e l v s h o r t p r e - b u r i a l r e s i c l e n c vo n t h e l l o ( ) d p l a i n . C h e r r i c a l e t c h i n g o f corticrl bonc rnd thorough disarticulati.n .f tlrc carc.sscsare taphonorric inciicaiors c,rrsistentrvith sedinrent'lrgical cvidence that the bones wcre <Jeposited i n t a a * e t . m u d d r ' . s u h t r . p i c a l e n v i r o n ' r e n t . I r u r t h e rc o r r p a r i s . n s r v i r h i n - c h a n n e l b o n e b e c l si n s l l n d s t o n c f a c i e s m u s t n o \ r b e u n d e r t . l k e nt { ) b c t t e r u r r r j e r s t l n c tl } r e p a l e o c ' c o l o l i c a lr e l a t i o n s h i p sb e t w e c n t h c s e d c p o s i t s . P R O T O . . \ V IA N i \ F F I N I T Y O F T H I 1 P L t , S| ( ) R N t S ' f l t A C K N . { AK E R C i l F - I t L I N S K I . G c r a r d . P o l . C e o l . I n s t . . I l a k o * i c c k a - 1 . W a r s a r v .{ X ) ( . } 7 5 P O L r \ N I ) : S A B A T H . K a r o l . I n s r . o i p a l e o b i o l o g r , .T r v a r i l a 5 l / - 5 - 5 . Warsas'. (X)-llI u PO[-AND. Despitc doubts cast ()lt sLrpposedprc-/\r( lu(()pter\..r clsteitlogical lnd some ich_ nological firssil record of birds. Pl<tiornis tracks display remarkably avian t'ea_ t u r e s .T h o s e d i s t i n c t i v e .s m a l l ( 3 _ 5 - - 5 n3 r n r l o n g ) f b o t p r i n r so c c u r i n t h e L . j .T r i a s s i c o f V i r l i n i a . I - . J u r a s s i co f M a s s a c h u s c t t sa n d p o l a n d . T h e t r a c k s s h a r et w . t e a t u r e s a b s e n l l r . n r n o n a v i a n d i n o s a u r t r a c k s : 1 1 ) b i r d l i k e r e r , e r s e dh a l l ' x : { l ) s r v e l l e d p r o x i m a l p a d o 1 - r h ed i g i t I I I . w j t h n o l a t e r a l r o e s ' p r o x i n r a l p a c l s . T h e P l a . t i o r n i . sa n d A r c . h u e o p r e t r- . rh a l l u r l e n g r h e q u u l r o n l y l / 3 o l t h e t h i r d d i g i t l e n g t h . w h e r e a s h a l l u x o f l a t e r b i r d s . e . g . S i / r . / r , r i . ri,s n e a r l y h a l f a s l o n g a s t h e t h i r d t o e . a n d i n s o m e m o d e r n l l v i n g b i r d s t h e s ed i g i r s a r e o f t h e s a n t e l e n g t h . The relersed avian hallux and swelled proximal pad ol digit lll. extentling the m a i n g r i p p i n g s u r f a c ea x i a l l y f r o r r t h e d i g i t I I I r o h a l l u x . e v i d e n t l y e n h a n c eg i a s p i n g a b i l i t ) , .T h i s f b o t a d a p t a t i o nm a y h a v e e v o l v e d p a r a l l e l t o w i n g s ' d e v e l o p m e n t . compensating tbr the deterioration of the forelimbs' grasping functions. Thus. if t h e h a l l u x e n l a r g e m e n ti n d e e d r e f l e c t sa n a d a p t a t i o n a lt r e n d o f b i r d f o o t e v o l u t i o n . and assuming a cursorial hypothesis firr the origin of birds. rhe pl"ri.)r,ri., trackmaker might even have been a fairlv advanccd flving creatcrre. 46A Plesi<trni,swouid imply emergence ot bircls as early as near the T-J bounclrry. before the Maniraptora appeared. Thus. their avian traits. includinr: sicklc-clau,ed p e d a l d i g i t I l - f o u n d r e c e n t l y i n r b i r c l R t r / r r a r z r .n a y h a v e b e e n i n h r , f i r c d l r ) n r their protoitvian ancestor."not vlce versa. EARLY EOCENE LAND MAMMALS FROM I-HE MTDDLE AND T]PPER GHAZIJ FORMATION OF BALUCHISTAN (PAKISTAN) C I N G E R I C H . P h i l i p D . , G e o l . S c i . l n d N l u s . P a l e o n r o l . .U n i r ' . N . l i c h i - r a n . A n n A r b o r . M I , 4 8 1 0 9 - 1 0 7 9 : A R I F . M u h r m r n a d .G e o l . S t r r v .p a k . . I s l a m a b a d :K H A N , I n r i z a r H u s s a i n . G e o l . S u r v . p a k . . e u e l r a : C L y D I l . W i l l i a m C . . E a r t h S c i e n c e s .L . J n i vN . c u . l . l a m p s h i r e .D u r h a l r . N H . 031324--3589 B:t - O C H . J o n i t r h a n1 . . C i e o l . S c i . a n c l N { u s . p r l e o n r o l . . U n i v . M i c h i g a n ,A n n A r b o n M l . + t 3 1 0 9 - 1 0 7 9 . T h e c o n d y l a r t h Q u e t u t t o t t l t u r a t h u i c a n t e l . n t n t t h c c o u l - b e a r i n gn t i c l c l l eC h a z i . j Formation of the Sor Range near Quetta. Tu'o (lueilirclonids trc n()\\. Kn()\\.nlronl middle Chazij coals near Mach. Thc uppcr Chazi.i Fornution netr euctll is il t h i c k s e q u e n c eo l r n u d s t ( ) n c - r i c hw i l l r v r r r c l - l i k e l l u ' i a l t l c p o s i t s r r i r h l i r n L l . n r c r a t e s a t t h c b a s e i n p l a c e s a n d r v c l l c l e v c l o p e crl c l l o v . r c c l . a n d p u r p l c p a l c o s o l s a b o v e . I t h a s y i c l d e d a d i v e r s i t v o l q u e t t a c _ r ' o n i da s.large tillodont. ilnd r huno_ l o p h o d o n t p e r i s s o d a c t y l .A t K i n g r i . s o n r e l 5 O k r r r r o r h e e a s l . r h e u p p c f C i h a z i .ijs m o r e s a n d r i c h a n d M u r r c e - i i k e . a n c l r v c l i r u n c l t i l l o d o n t s . p e r i s s o r i a c t v l sr.n r l l r r t h r a c o b u n i d s .b u t n o q u e t t . c y o n i d s . F . r a n r i n i l c r - r l l 1 ) n r u n d e r l n t l . r ' c r l r i n g r ' . r n l a t i o n s i n d i c a t e t h a t G h a z i . in r a m m a l s i n I l a l u c h i s l u n c o n r e l r r n r p l u n k l t r n i c z o r c P 6 a n d s h a l l o w b e n t h i c z o n e s S B f l l { } t n t i t i - \ , p f e s i a n ) . K L r l d i l n u - S u b ; i 1 hlrar' c l m a n r m a l s f r o t n l ) u k i s l u r ra n r l I n d i a c o r n c l i r n t z o n c s P 9 l 0 a n d S B I I l . t r t , r r r l r m i d d l e E . c e n e ) a . ( l r c p r c s e n .t I a l c r t a r r n i (r l o r r i r r r t c dh r r r t i i , t l u e rlr. I l r r r i h . r , r l F o r n r a t i o nw a s s h c c lc i t s t u a r d t i l r r i r r t l l n t i r , l ' . r k i s t . r nt i , n r r r n o l l s h o r c i : l r r r r l e h a i n : t l s n r o s t c o m p l e t e p r o x i m a l e x p r c s s i o nn L N ' b c ' i r r H r l a r i l \ t r r t r I l a P p e ( l i l s \ l l r r f c c F o r n r a t i o nt h a t s p l r n S I 3 8 t h r o u g h S B l l - . l l . S A U R O P O D C I I A N I A L E L E M F - N ' l S F I t O N . t N I , , \ l _ A \ \ , t : t i \ i l ) t _ t,(\ - t - l ( ) \ s ( ) l : TITANOSAURID CRANIAL NiOItPHOI-(XJY G O M A N I . E l i z a b e t h M . . D e p a r t m e n ro i C j e o l o s i c a lS c i e n c c s .S h u l c r M u s e u n r . S o u r h e r n M e r h o c l i \ t L J l i v c rs i r v - D a l l a s . ' f X . 7 5 1 7 - i . t , S \ . T i t a n o s a u r i d s h a v c b e e n c o n s i c l e r e da s h a v i n g l ) i p l t i l o t L r . s - l i k es k u l l s . i n c c r h e , c r a n i u m o f A n t ( r . t o . t ( ! r r t ! . \ , a t i l l l t O s l u r i t i - \ \ , l s f c c o n s t r u c t c dr r i t h l t n c l o n L a t c t l p r e o r b i t a l r e g i o n . H O r v e v e r .r e c e n t s l u d i c s s u - q c e stth r t t h e c [ r n i a o l ' t i l r r i r s u r r r i t l s a r e h i g h a n d s h o r t a s i n B r u c l t i o s a u n r . ra n c l( t t t u t r n r . r t t n r C . r l.a c l i s t i cr n a l r s c : h r r . , b e e n o f l i t t l e h e l p i n s o l v i n g t h e p r o b l c n t o l t i r u n o s r u r i d c r . u n i . r rl n , , r l , h , , l , , l r l . r c a u s e t i t a n o s a u r i d sh r v e o f l e n b e e n r e p r e s e t r t c da s b e i n l c l o s c l y r e l i l t c ( l t ( ) c i t h c f c l i p l o d o c . i d s o r b r a c h i o s a u r i d sT . h e r e i i r r c .r h e ( ) c c L r r r e n c.cf a b r s i c r a r i r r r r . r l u a r l r a t e . J u g a l , p a r t i a l l r a x i l l a . p r e m a x i l l a . p t e r v - s o i d s c. c t o p t e r v L o i c l .c l c n r a r r .a n d I s p f e n i a l o f M a l u t i . s t r r r u . s ,a t i t a n o s r u r i d . r n c l i t c l e n t l r v . 1 . s n r t l l c r t a r . n l a r n r M a l a w i i s s i g n i l i c a n t .T h i s m a t e r i r l \ u g g c s r \ r h r t t i r r n r i s r u r i t l sh a \ c l c s : f c l f i l c r e ( l n a s a l s t h a n D i p l o d o c u . s ,t h e r o s t r u t n i s s h o r t . l l r e c r r n i u n t i s h i ! h . t h c r ( ) ( ) l hr ( r * i s e i t h e r a n t e r i o r l y f e s t r i c t c do r c x t c n ( l \ i r l o r r gh r l l t h c r l e n l a r r . i r n r l t l L r t l c c t h l r r e primitivelv slightly brorcl or cvlinclricrl. C A R B O N A N D O X ) ' C I r N I S O T O P I T SI N - f ( l O t - t I t i N i \ i \ i l : l _ O l j t : O ( - t : \ t : L A N D M A M M A T - S A N D A R C H / \ E O C F _ 1 ' t rw H A t _ E S C O S W A M I . A n . j a l i .D e p t . C l e o l o s i c aSl c i c n c c s t. . J n i l c r \ i r vo l N l i c h i u r r n . A n n A r b o r _ M I . ; t r i 3 1 0 9(:I I N C E R I C t { . P h i l i p l ) . . D c p r . C i c o t o r i r . i r l S c i e n c c sa n t i M u s . P a l e o n r o l o g r ' t- J n i v c r s i t l -o l l \ , 1 i c h i g a n,.\ n n \ r b ( ) r . M I . . 1 8t { X ) t 0 7 9 . I s o t o p i c a n a l y s i s o l p h o s p h a t eh a s b e e n u s c c l1 o i n l c r o s n r r r c g r r l l t o r r a r l l P t a r i o n to ingestion ol' seit wirter early in *.hrlc c'.lrrti.'. \\t rriet{ l. cxlc.ri lhi: br n t e a s u r i n gt h e i s o t o l t i c c o n t p o s i t i o n o t c l r b r r n r n d o x \ ! c n i l e n . r r l e l e l , l , , , n l r t . . a n d p h o s p h a t ei n r * i c l c r r l l n r e o l - t c r r e s t r i a ln r i r n r r n l l s .t r a n s i t ( r n r r r r c h i r c ( ) . c r c s . a r r c lm l r i n c a r c h a c o c c t e sS. ' O ( S M O W ) l r 1 ) n ln l r ( ) s l ) h a t rca n s e s l r r r n t 1 6 . - 5r L rl l . l p e r n r i l f o r t e f r e s l r i t l K l t i r t l u r i u i r n c lL L t u t t t t t l l t r t t t i rfrr.o r t r l ( r . J t ( ) l 0 - i l i r l { r r l t s i t i o n a l P o A r ' c c l r r .nrn t l 1 ) c t l u t t i . s r t 'r.nsd. l i i . + t o l i ) . - i l i r r n l r r i n c / l r ) r l l / . ) r . . , r r lB\ .t l , i , / u t t ' t u s , D o r u d r t n , L \ t 1 dB o s i l ( ) . \ u u n t . \E. r r C i i ) l ) B ) l 1 1 ) n t c r r b ( ) n r t c r a n g c c l r . ( ) n r S . 9 to 8.2 for Kltirthuriu and ltutrrnitlltuuirr. l -5.0 to -, I I.-l lirr prr(rrllrir. ij.+ fol Dalurtiste.s. and 8.5 to -6.13 ft>r Rotllutt,trt.s. Iltbittt t,ttt.:.Donrdort. anrl grrrrlrr s a a n r s . 6 r N Od o e s n o t d i s t i n g u i s h l a n c l . t r r n s i l i o r r a l . o r n a r i n c n r a n t n r r l : i n o u r s a m p l e .L i g h t e r b r r C c l i s t i n g u i s l r eP s u k i t t ' t t t . rs' r . r r l l l l r n d . t r i i r s i t i o n a l .r n c l ' r r i n e n r a m m a l s s t u d i e d h c r e . L i g h t e r E r r c ' a l t r e s g c n e r a l l v c o r r e l a t er r i t b r r r - r r c s r r i r l d i e t , b r r t h e a v y 6 r ' C v a l u e s o b t a i n e r !I o r K l r i i l r , u i , t L r ' n Lt lt t t t n t i t l l t u r t i l / . ( ) 4 r p l i c a r a s i m p l e i n t e r p r e t a t i o n .O x y g e n i s o t o p i c c ( ) n t p o s i t i o ni s i n f l u e t r c c , cblv p o o r l r r r n d e r s t o o d m e t a b o l i c f a c t o r s . s o o b s e r v e c l' a r i a t i o n i s p r e s e n t l vr l i f t i c u l t r o a t t r i h r r t c to any sinrplechange in environmenl or 1th1'siolouv. THE GREAT-TOOI-I-IED LAMN I D S I-lA lt KS : Cr1RCHA I?OD()N, NO't' CARCHAROCLES G O T T F R I E D . M i c h a e l D . . M S U M u s c u n r . M i c h i - c a nS t a r c U n i r , . . H a s r Lansing. MI. 48824,1045. USA. The fbssil 'great-toothed' lantnid sharks havc been assignetl by sontc ro (.ru. c h u x t d o r t s m i t h . l l t 3 l t ( w h i c h i n c l u d e sr h c c r r a n r G r e a r w h i r c S h a r k ( i r r r l r r r r r r r l r r i O 1998 by the Society of VertebratePaleontologl'